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    Gerald Kooyman

    TitleEmeritus Professor
    SchoolUniversity of California, San Diego
    DepartmentScripps Institution of Oceanography Department
    Address9500 Gilman Drive #0204
    CA La Jolla 92093
    Phone858-534-2091
    vCardDownload vCard

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      Publications listed below are automatically derived from MEDLINE/PubMed and other sources, which might result in incorrect or missing publications. Researchers can login to make corrections and additions, or contact us for help.
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      1. Hays GC, Ferreira LC, Sequeira AM, Meekan MG, Duarte CM, Bailey H, Bailleul F, Bowen WD, Caley MJ, Costa DP, Eguíluz VM, Fossette S, Friedlaender AS, Gales N, Gleiss AC, Gunn J, Harcourt R, Hazen EL, Heithaus MR, Heupel M, Holland K, Horning M, Jonsen I, Kooyman G, Lowe CG, Madsen PT, Marsh H, Phillips RA, Righton D, Ropert-Coudert Y, Sato K, Shaffer SA, Simpfendorfer CA, Sims DW, Skomal G, Takahashi A, Trathan PN, Wikelski M, Womble JN, Thums M. Key Questions in Marine Megafauna Movement Ecology. Trends Ecol Evol. 2016 Jun; 31(6):463-75. PMID: 26979550.
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      2. Williams CL, Hagelin JC, Kooyman G. Hidden keys to survival: the type, density, pattern and functional role of emperor penguin body feathers. Proc Biol Sci. 2015 Oct 22; 282(1817). PMID: 26490794.
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      3. Younger JL, Clucas GV, Kooyman G, Wienecke B, Rogers AD, Trathan PN, Hart T, Miller KJ. Too much of a good thing: sea ice extent may have forced emperor penguins into refugia during the last glacial maximum. Glob Chang Biol. 2015 Jun; 21(6):2215-26. PMID: 25728986.
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      4. Kooyman G. Marine mammals and Emperor penguins: a few applications of the Krogh principle. Am J Physiol Regul Integr Comp Physiol. 2015 Jan 15; 308(2):R96-104. PMID: 25411360.
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      5. Fretwell PT, Trathan PN, Wienecke B, Kooyman G. Emperor penguins breeding on iceshelves. PLoS One. 2014; 9(1):e85285. PMID: 24416381.
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      6. Meir JU, Robinson PW, Vilchis LI, Kooyman G, Costa DP, Ponganis PJ. Blood oxygen depletion is independent of dive function in a deep diving vertebrate, the northern elephant seal. PLoS One. 2013; 8(12):e83248. PMID: 24376671.
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      7. Fretwell PT, Larue MA, Morin P, Kooyman G, Wienecke B, Ratcliffe N, Fox AJ, Fleming AH, Porter C, Trathan PN. An emperor penguin population estimate: the first global, synoptic survey of a species from space. PLoS One. 2012; 7(4):e33751. PMID: 22514609.
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      8. Sato K, Shiomi K, Marshall G, Kooyman G, Ponganis PJ. Stroke rates and diving air volumes of emperor penguins: implications for dive performance. J Exp Biol. 2011 Sep 1; 214(Pt 17):2854-63. PMID: 21832128.
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      9. Blight LK, Ainley DG, Ackley SF, Ballard G, Ballerini T, Brownell RL, Cheng CH, Chiantore M, Costa D, Coulter MC, Dayton P, Devries AL, Dunbar R, Earle S, Eastman JT, Emslie SD, Evans CW, Garrott RA, Kim S, Kooyman G, Lescroël A, Lizotte M, Massaro M, Olmastroni S, Ponganis PJ, Russell J, Siniff DB, Smith WO, Stewart BS, Stirling I, Willis J, Wilson P, Woehler EJ. Fishing for data in the Ross Sea. Science. 2010 Dec 3; 330(6009):1316. PMID: 21127229.
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      10. Kooyman G. First in the measure of energetics in a swimming tetrapod. J Exp Biol. 2010 May; 213(Pt 10):1609-10. PMID: 20435809.
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      11. Jobsis PD, Ponganis PJ, Kooyman G. Effects of training on forced submersion responses in harbor seals. J Exp Biol. 2001 Nov; 204(Pt 22):3877-85. PMID: 11807105.
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      12. Nagy KA, Kooyman G, Ponganis PJ. Energetic cost of foraging in free-diving emperor penguins. Physiol Biochem Zool. 2001 Jul-Aug; 74(4):541-7. PMID: 11436138.
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      13. Trillmich F, Kooyman G. Field metabolic rate of lactating female Galápagos fur seals (Arctocephalus galapagoensis): the influence of offspring age and environment. Comp Biochem Physiol A Mol Integr Physiol. 2001 Jul; 129(4):741-9. PMID: 11440861.
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      14. Ancel A, Starke LN, Ponganis PJ, Van Dam R, Kooyman G. Energetics of surface swimming in Brandt's cormorants (Phalacrocorax penicillatus brandt). J Exp Biol. 2000 Dec; 203(Pt 24):3727-31. PMID: 11076736.
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      15. Ponganis PJ, Kooyman G. Diving physiology of birds: a history of studies on polar species. Comp Biochem Physiol A Mol Integr Physiol. 2000 Jun; 126(2):143-51. PMID: 10936756.
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      16. Ponganis PJ, Starke LN, Horning M, Kooyman G. Development of diving capacity in emperor penguins. J Exp Biol. 1999 Apr; 202(Pt 7):781-6. PMID: 10069967.
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      17. Dolar ML, Suarez P, Ponganis PJ, Kooyman G. Myoglobin in pelagic small cetaceans. J Exp Biol. 1999 Feb; 202(Pt 3):227-36. PMID: 9882635.
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      18. Kooyman G, Ponganis PJ. The physiological basis of diving to depth: birds and mammals. Annu Rev Physiol. 1998; 60:19-32. PMID: 9558452.
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      19. Ponganis PJ, Costello ML, Starke LN, Mathieu-Costello O, Kooyman G. Structural and biochemical characteristics of locomotory muscles of emperor penguins, Aptenodytes forsteri. Respir Physiol. 1997 Jul; 109(1):73-80. PMID: 9271809.
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      20. Ponganis PJ, Kooyman G, Starke LN, Kooyman CA, Kooyman TG. Post-dive blood lactate concentrations in emperor penguins, Aptenodytes forsteri. J Exp Biol. 1997 Jun; 200(Pt 11):1623-6. PMID: 9202449.
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      21. Ponganis PJ, Kooyman G, Winter LM, Starke LN. Heart rate and plasma lactate responses during submerged swimming and trained diving in California sea lions, Zalophus californianus. J Comp Physiol B. 1997 Jan; 167(1):9-16. PMID: 9051904.
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      22. Ancel A, Horning M, Kooyman G. Prey ingestion revealed by oesophagus and stomach temperature recordings in cormorants. J Exp Biol. 1997 Jan; 200(Pt 1):149-54. PMID: 9023995.
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      23. Kooyman G, Ponganis PJ. Emperor penguin oxygen consumption, heart rate and plasma lactate levels during graded swimming exercise. J Exp Biol. 1994 Oct; 195:199-209. PMID: 7964411.
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      24. Ponganis PJ, Kooyman G, Castellini MA, Ponganis EP, Ponganis KV. Muscle temperature and swim velocity profiles during diving in a Weddell seal, Leptonychotes weddellii. J Exp Biol. 1993 Oct; 183:341-6. PMID: 8245765.
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      25. Castellini MA, Kooyman G, Ponganis PJ. Metabolic rates of freely diving Weddell seals: correlations with oxygen stores, swim velocity and diving duration. J Exp Biol. 1992 Apr; 165:181-94. PMID: 1588250.
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      26. Kooyman G, Ponganis PJ, Castellini MA, Ponganis EP, Ponganis KV, Thorson PH, Eckert SA, LeMaho Y. Heart rates and swim speeds of emperor penguins diving under sea ice. J Exp Biol. 1992 Apr; 165:161-80. PMID: 1588249.
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      27. Ponganis PJ, Kooyman G, Sartoris D, Jobsis P. Pinniped splenic volumes. Am J Physiol. 1992 Feb; 262(2 Pt 2):R322-5. PMID: 1539742.
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      28. Davis RW, Castellini MA, Williams TM, Kooyman G. Fuel homeostasis in the harbor seal during submerged swimming. J Comp Physiol B. 1991; 160(6):627-35. PMID: 2045543.
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      29. Williams TM, Kooyman G, Croll DA. The effect of submergence on heart rate and oxygen consumption of swimming seals and sea lions. J Comp Physiol B. 1991; 160(6):637-44. PMID: 2045544.
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      30. Ponganis PJ, Kooyman G, Zornow MH, Castellini MA, Croll DA. Cardiac output and stroke volume in swimming harbor seals. J Comp Physiol B. 1990; 160(5):473-82. PMID: 2097298.
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      31. Castellini MA, Kooyman G. Behavior of freely diving animals. Undersea Biomed Res. 1989 Sep; 16(5):355-62. PMID: 2678666.
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      32. Davis RW, Castellini MA, Kooyman G, Maue R. Renal glomerular filtration rate and hepatic blood flow during voluntary diving in Weddell seals. Am J Physiol. 1983 Nov; 245(5 Pt 1):R743-8. PMID: 6638219.
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      33. Drabek CM, Kooyman G. Terminal airway embryology of the delphinid porpoises, Stenella attenuata and S. longirostris. J Morphol. 1983 Jan; 175(1):65-72. PMID: 6842612.
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      34. Kooyman G, Davis RW, Croxall JP, Costa DP. Diving depths and energy requirements of king penguins. Science. 1982 Aug 20; 217(4561):726-7. PMID: 7100916.
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      35. Sinnett EE, Wahrenbrock EA, Kooyman G. Cardiovascular depression and thermoregulatory disruption caused by pentothal/halothane anesthesia in the harbor seal, Phoca vitulina. J Wildl Dis. 1981 Jan; 17(1):121-30. PMID: 7253094.
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      36. Kooyman G, Castellini MA, Davis RW. Physiology of diving in marine mammals. Annu Rev Physiol. 1981; 43:343-56. PMID: 7011189.
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      37. Kooyman G, Sinnett EE. Mechanical properties of the harbor porpoise lung, Phocoena phocoena. Respir Physiol. 1979 Apr; 36(3):287-300. PMID: 441581.
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      38. Sinnett EE, Kooyman G, Wahrenbrock EA. Pulmonary circulation of the harbor seal. J Appl Physiol Respir Environ Exerc Physiol. 1978 Nov; 45(5):718-27. PMID: 730568.
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      39. Kooyman G, Gentry RL, Urguhart DL. Northern fur seal diving behavior: a new approach to its study. Science. 1976 Jul 30; 193(4251):411-2. PMID: 935876.
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      40. Kooyman G, Gentry RL, Bergman WP, Hammel HT. Heat loss in penguins during immersion and compression. Comp Biochem Physiol A Comp Physiol. 1976; 54(1):75-80. PMID: 3348.
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      41. Kooyman G, Schroeder JP, Greene DG, Smith VA. Gas exchange in penguins during simulated dives to 30 and 68 m. Am J Physiol. 1973 Dec; 225(6):1467-71. PMID: 4760462.
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      42. Kooyman G, Kerem DH, Campbell WB, Wright JJ. Pulmonary gas exchange in freely diving Weddell seals, Leptonychotes weddelli. Respir Physiol. 1973 Apr; 17(3):283-90. PMID: 4702979.
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      43. Denison DM, Kooyman G. The structure and function of the small airways in pinniped and sea otter lungs. Respir Physiol. 1973 Jan; 17(1):1-10. PMID: 4688284.
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      44. Kooyman G, Schroeder JP, Denison DM, Hammond DD, Wright JJ, Bergman WP. Blood nitrogen tensions of seals during simulated deep dives. Am J Physiol. 1972 Nov; 223(5):1016-20. PMID: 4654334.
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      45. Kooyman G, Campbell WB. Heart rates in freely diving Weddell Seals, Leptonychotes weddelli. Comp Biochem Physiol A Comp Physiol. 1972 Sep 1; 43(1):31-6. PMID: 4404585.
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      46. Kerem D, Kooyman G, Schroeder JP, Wright JJ, Drabek CM. Hyperbaric oxygen-induced seizure in a marine mammal, the seal. Am J Physiol. 1972 May; 222(5):1322-5. PMID: 5022390.
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      47. Kooyman G. Deep diving behaviour and effects of pressure in reptiles, birds, and mammals. Symp Soc Exp Biol. 1972; 26:295-311. PMID: 4574858.
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      48. Kooyman G, Kerem DH, Campbell WB, Wright JJ. Pulmonary function in freely diving Weddell seals, Leptonychotes weddelli. Respir Physiol. 1971 Aug; 12(3):271-82. PMID: 5136018.
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      49. Kooyman G, Hammond DD, Schroeder JP. Bronchograms and tracheograms of seals under pressure. Science. 1970 Jul 3; 169(3940):82-4. PMID: 5447537.
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      50. Kooyman G. The Weddell seal. Sci Am. 1969 Aug; 221(2):101-6. PMID: 5791821.
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      51. Lenfant C, Elsner R, Kooyman G, Drabek CM. Respiratory function of blood of the adult and fetus Weddell seal Leptonychotes weddelli. Am J Physiol. 1969 Jun; 216(6):1595-7. PMID: 5786750.
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      52. Lenfant C, Kooyman G, Elsner R, Drabek CM. Respiratory function of blood of the Adélie penguin Pygoscelis adeliae. Am J Physiol. 1969 Jun; 216(6):1598-600. PMID: 5786751.
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      53. Kooyman G. Maximum diving capacities of the Weddell seal, Leptonychotes weddelli. Science. 1966 Mar 25; 151(3717):1553-4. PMID: 5909593.
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