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    Robert Shadwick

    TitleEmeritus Professor
    SchoolUniversity of California, San Diego
    DepartmentScripps Institution of Oceanography Department
    Address9500 Gilman Drive #0202
    CA La Jolla 92093
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      Publications listed below are automatically derived from MEDLINE/PubMed and other sources, which might result in incorrect or missing publications. Researchers can login to make corrections and additions, or contact us for help.
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      1. Lillie MA, Vogl AW, Gil KN, Gosline JM, Shadwick R. Two Levels of Waviness Are Necessary to Package the Highly Extensible Nerves in Rorqual Whales. Curr Biol. 2017 Mar 06; 27(5):673-679. PMID: 28216319.
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      2. Morell M, Brownlow A, McGovern B, Raverty SA, Shadwick R, André M. Implementation of a method to visualize noise-induced hearing loss in mass stranded cetaceans. Sci Rep. 2017 Feb 06; 7:41848. PMID: 28165504.
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      3. Shadwick R, Denny MW. John Moffit Gosline, BA, PhD, FRSC (1943-2016). J Exp Biol. 2017 Feb 01; 220(Pt 3):334-335. PMID: 28148816.
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      4. Werth AJ, Straley JM, Shadwick R. Baleen wear reveals intraoral water flow patterns of mysticete filter feeding. J Morphol. 2016 Apr; 277(4):453-71. PMID: 26825852.
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      5. Clark TE, Lillie MA, Vogl AW, Gosline JM, Shadwick R. Mechanical contribution of lamellar and interlamellar elastin along the mouse aorta. J Biomech. 2015 Oct 15; 48(13):3599-605. PMID: 26321365.
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      6. Vogl AW, Lillie MA, Piscitelli MA, Goldbogen JA, Pyenson ND, Shadwick R. Stretchy nerves are an essential component of the extreme feeding mechanism of rorqual whales. Curr Biol. 2015 May 04; 25(9):R360-1. PMID: 25942546.
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      7. Morell M, Lenoir M, Shadwick R, Jauniaux T, Dabin W, Begeman L, Ferreira M, Maestre I, Degollada E, Hernandez-Milian G, Cazevieille C, Fortuño JM, Vogl W, Puel JL, André M. Ultrastructure of the Odontocete organ of Corti: scanning and transmission electron microscopy. J Comp Neurol. 2015 Feb 15; 523(3):431-48. PMID: 25269663.
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      8. Piscitelli MA, Raverty SA, Lillie MA, Shadwick R. A review of cetacean lung morphology and mechanics. J Morphol. 2013 Dec; 274(12):1425-40. PMID: 24027086.
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      9. Lillie MA, Piscitelli MA, Vogl AW, Gosline JM, Shadwick R. Cardiovascular design in fin whales: high-stiffness arteries protect against adverse pressure gradients at depth. J Exp Biol. 2013 Jul 15; 216(Pt 14):2548-63. PMID: 23804669.
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      10. Pinto SJ, Shadwick R. Material and structural properties of fin whale (Balaenoptera physalus) Zwischensubstanz. J Morphol. 2013 Aug; 274(8):947-55. PMID: 23640788.
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      11. Shadwick R, Goldbogen JA, Potvin J, Pyenson ND, Vogl AW. Novel muscle and connective tissue design enables high extensibility and controls engulfment volume in lunge-feeding rorqual whales. J Exp Biol. 2013 Jul 15; 216(Pt 14):2691-701. PMID: 23580724.
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      12. Potvin J, Goldbogen JA, Shadwick R. Metabolic expenditures of lunge feeding rorquals across scale: implications for the evolution of filter feeding and the limits to maximum body size. PLoS One. 2012; 7(9):e44854. PMID: 23024769; PMCID: PMC3443106.
        View in: PubMed, PubMed Central
      13. Lillie MA, Armstrong TE, Gérard SG, Shadwick R, Gosline JM. Contribution of elastin and collagen to the inflation response of the pig thoracic aorta: assessing elastin's role in mechanical homeostasis. J Biomech. 2012 Aug 09; 45(12):2133-41. PMID: 22770359.
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      14. Pyenson ND, Goldbogen JA, Vogl AW, Szathmary G, Drake RL, Shadwick R. Discovery of a sensory organ that coordinates lunge feeding in rorqual whales. Nature. 2012 May 23; 485(7399):498-501. PMID: 22622577.
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      15. Shadwick R, Goldbogen JA. Muscle function and swimming in sharks. J Fish Biol. 2012 Apr; 80(5):1904-39. PMID: 22497412.
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      16. Schilt CR, Cranford TW, Krysl P, Shadwick R, Hawkins AD. Vibration of the otoliths in a teleost. Adv Exp Med Biol. 2012; 730:105-7. PMID: 22278460.
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      17. Syme DA, Shadwick R. Red muscle function in stiff-bodied swimmers: there and almost back again. Philos Trans R Soc Lond B Biol Sci. 2011 May 27; 366(1570):1507-15. PMID: 21502122; PMCID: PMC3130444.
        View in: PubMed, PubMed Central
      18. Field DJ, Lin SC, Ben-Zvi M, Goldbogen JA, Shadwick R. Convergent evolution driven by similar feeding mechanics in balaenopterid whales and pelicans. Anat Rec (Hoboken). 2011 Aug; 294(8):1273-82. PMID: 21618438.
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      19. Goldbogen JA, Calambokidis J, Oleson E, Potvin J, Pyenson ND, Schorr G, Shadwick R. Mechanics, hydrodynamics and energetics of blue whale lunge feeding: efficiency dependence on krill density. J Exp Biol. 2011 Jan 01; 214(Pt 1):131-46. PMID: 21147977.
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      20. Potvin J, Goldbogen JA, Shadwick R. Scaling of lunge feeding in rorqual whales: an integrated model of engulfment duration. J Theor Biol. 2010 Dec 07; 267(3):437-53. PMID: 20816685.
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      21. Field DJ, Campbell-Malone R, Goldbogen JA, Shadwick R. Quantitative computed tomography of humpback whale (Megaptera novaeangliae) mandibles: mechanical implications for rorqual lunge-feeding. Anat Rec (Hoboken). 2010 Jul; 293(7):1240-7. PMID: 20583267.
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      22. Lillie MA, Shadwick R, Gosline JM. Mechanical anisotropy of inflated elastic tissue from the pig aorta. J Biomech. 2010 Aug 10; 43(11):2070-8. PMID: 20430395.
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      23. Goldbogen JA, Potvin J, Shadwick R. Skull and buccal cavity allometry increase mass-specific engulfment capacity in fin whales. Proc Biol Sci. 2010 Mar 22; 277(1683):861-8. PMID: 19939846; PMCID: PMC2842724.
        View in: PubMed, PubMed Central
      24. Potvin J, Goldbogen JA, Shadwick R. Passive versus active engulfment: verdict from trajectory simulations of lunge-feeding fin whales Balaenoptera physalus. J R Soc Interface. 2009 Nov 06; 6(40):1005-25. PMID: 19158011; PMCID: PMC2827442.
        View in: PubMed, PubMed Central
      25. Goldbogen JA, Calambokidis J, Croll DA, Harvey JT, Newton KM, Oleson EM, Schorr G, Shadwick R. Foraging behavior of humpback whales: kinematic and respiratory patterns suggest a high cost for a lunge. J Exp Biol. 2008 Dec; 211(Pt 23):3712-9. PMID: 19011211.
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      26. Shadwick R, Syme DA. Thunniform swimming: muscle dynamics and mechanical power production of aerobic fibres in yellowfin tuna (Thunnus albacares). J Exp Biol. 2008 May; 211(Pt 10):1603-11. PMID: 18456888.
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      27. Cranford TW, McKenna MF, Soldevilla MS, Wiggins SM, Goldbogen JA, Shadwick R, Krysl P, St Leger JA, Hildebrand JA. Anatomic geometry of sound transmission and reception in Cuvier's beaked whale (Ziphius cavirostris). Anat Rec (Hoboken). 2008 Apr; 291(4):353-78. PMID: 18228579.
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      28. Shadwick R. Foundations of animal hydraulics: geodesic fibres control the shape of soft bodied animals. J Exp Biol. 2008 Feb; 211(Pt 3):289-91. PMID: 18203982.
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      29. Donley JM, Shadwick R, Sepulveda CA, Syme DA. Thermal dependence of contractile properties of the aerobic locomotor muscle in the leopard shark and shortfin mako shark. J Exp Biol. 2007 Apr; 210(Pt 7):1194-203. PMID: 17371918.
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      30. Rapoport HS, Shadwick R. Reversibly labile, sclerotization-induced elastic properties in a keratin analog from marine snails: whelk egg capsule biopolymer (WECB). J Exp Biol. 2007 Jan; 210(Pt 1):12-26. PMID: 17170144.
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      31. Gemballa S, Konstantinidis P, Donley JM, Sepulveda C, Shadwick R. Evolution of high-performance swimming in sharks: transformations of the musculotendinous system from subcarangiform to thunniform swimmers. J Morphol. 2006 Apr; 267(4):477-93. PMID: 16429422.
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      32. Goldbogen JA, Calambokidis J, Shadwick R, Oleson EM, McDonald MA, Hildebrand JA. Kinematics of foraging dives and lunge-feeding in fin whales. J Exp Biol. 2006 Apr; 209(Pt 7):1231-44. PMID: 16547295.
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      33. Bernal D, Donley JM, Shadwick R, Syme DA. Mammal-like muscles power swimming in a cold-water shark. Nature. 2005 Oct 27; 437(7063):1349-52. PMID: 16251963.
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      34. Soldevilla MS, McKenna MF, Wiggins SM, Shadwick R, Cranford TW, Hildebrand JA. Cuvier's beaked whale (Ziphius cavirostris) head tissues: physical properties and CT imaging. J Exp Biol. 2005 Jun; 208(Pt 12):2319-32. PMID: 15939773.
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      35. Donley JM, Shadwick R, Sepulveda CA, Konstantinidis P, Gemballa S. Patterns of red muscle strain/activation and body kinematics during steady swimming in a lamnid shark, the shortfin mako (Isurus oxyrinchus). J Exp Biol. 2005 Jun; 208(Pt 12):2377-87. PMID: 15939777.
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      36. Goldbogen JA, Shadwick R, Fudge DS, Gosline JM. Fast-start muscle dynamics in the rainbow trout Oncorhynchus mykiss: phase relationship of white muscle shortening and body curvature. J Exp Biol. 2005 Mar; 208(Pt 5):929-38. PMID: 15755891.
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      37. Donley JM, Sepulveda CA, Konstantinidis P, Gemballa S, Shadwick R. Convergent evolution in mechanical design of lamnid sharks and tunas. Nature. 2004 May 06; 429(6987):61-5. PMID: 15129279.
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      38. Donley JM, Shadwick R. Steady swimming muscle dynamics in the leopard shark Triakis semifasciata. J Exp Biol. 2003 Apr; 206(Pt 7):1117-26. PMID: 12604571.
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      39. Shadwick R, Rapoport HS, Fenger JM. Structure and function of tuna tail tendons. Comp Biochem Physiol A Mol Integr Physiol. 2002 Dec; 133(4):1109-25. PMID: 12485695.
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      40. Rapoport HS, Shadwick R. Mechanical characterization of an unusual elastic biomaterial from the egg capsules of marine snails (Busycon spp.). Biomacromolecules. 2002 Jan-Feb; 3(1):42-50. PMID: 11866554.
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      41. Syme DA, Shadwick R. Effects of longitudinal body position and swimming speed on mechanical power of deep red muscle from skipjack tuna (Katsuwonus pelamis). J Exp Biol. 2002 Jan; 205(Pt 2):189-200. PMID: 11821485.
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      42. Bernal D, Dickson KA, Shadwick R, Graham JB. Review: Analysis of the evolutionary convergence for high performance swimming in lamnid sharks and tunas. Comp Biochem Physiol A Mol Integr Physiol. 2001 Jun; 129(2-3):695-726. PMID: 11423338.
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      43. Nauen JC, Shadwick R. The dynamics and scaling of force production during the tail-flip escape response of the California spiny lobster Panulirus interruptus. J Exp Biol. 2001 May; 204(Pt 10):1817-30. PMID: 11316502.
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      44. Katz SL, Syme DA, Shadwick R. High-speed swimming. Enhanced power in yellowfin tuna. Nature. 2001 Apr 12; 410(6830):770-1. PMID: 11298434.
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      45. Szulgit GK, Shadwick R. Dynamic mechanical characterization of a mutable collagenous tissue: response of sea cucumber dermis to cell lysis and dermal extracts. J Exp Biol. 2000 May; 203(Pt 10):1539-50. PMID: 10769216.
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      46. Shadwick R. Mechanical design in arteries. J Exp Biol. 1999 Dec; 202(Pt 23):3305-13. PMID: 10562513.
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      47. Shadwick R, Katz SL, Korsmeyer KE, Knower T, Covell JW. Muscle dynamics in skipjack tuna: timing of red muscle shortening in relation to activation and body curvature during steady swimming. J Exp Biol. 1999; 202(Pt 16):2139-2150. PMID: 10409485.
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      48. Korsmeyer KE, Lai NC, Shadwick R, Graham JB. Heart rate and stroke volume contribution to cardiac output in swimming yellowfin tuna: response to exercise and temperature. J Exp Biol. 1997 Jul; 200(Pt 14):1975-86. PMID: 9246781.
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      49. Korsmeyer KE, Lai NC, Shadwick R, Graham JB. Oxygen transport and cardiovascular responses to exercise in the yellowfin tuna Thunnus albacares. J Exp Biol. 1997 Jul; 200(Pt 14):1987-97. PMID: 9246782.
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      50. Gosline JM, Shadwick R. The mechanical properties of fin whale arteries are explained by novel connective tissue designs. J Exp Biol. 1996 Apr; 199(Pt 4):985-97. PMID: 8788091.
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      51. Shadwick R, Gosline JM. Arterial Windkessels in marine mammals. Symp Soc Exp Biol. 1995; 49:243-52. PMID: 8571227.
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      52. Shadwick R, Gosline JM. Arterial mechanics in the fin whale suggest a unique hemodynamic design. Am J Physiol. 1994 Sep; 267(3 Pt 2):R805-18. PMID: 8092327.
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      53. Pollock CM, Shadwick R. Allometry of muscle, tendon, and elastic energy storage capacity in mammals. Am J Physiol. 1994 Mar; 266(3 Pt 2):R1022-31. PMID: 8160851.
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      54. Pollock CM, Shadwick R. Relationship between body mass and biomechanical properties of limb tendons in adult mammals. Am J Physiol. 1994 Mar; 266(3 Pt 2):R1016-21. PMID: 8160850.
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      55. Shadwick R, Russell AP, Lauff RF. The structure and mechanical design of rhinoceros dermal armour. Philos Trans R Soc Lond B Biol Sci. 1992 Sep 29; 337(1282):419-28. PMID: 1359589.
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      56. Griffiths RI, Shadwick R, Berger PJ. Functional importance of a highly elastic ligament on the mammalian diaphragm. Proc Biol Sci. 1992 Aug 22; 249(1325):199-204. PMID: 1360681.
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      57. Shadwick R. Elastic energy storage in tendons: mechanical differences related to function and age. J Appl Physiol (1985). 1990 Mar; 68(3):1033-40. PMID: 2341331.
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      58. Gibbons CA, Shadwick R. Functional similarities in the mechanical design of the aorta in lower vertebrates and mammals. Experientia. 1989 Dec 01; 45(11-12):1083-8. PMID: 2513219.
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      59. Shadwick R, Gosline JM. Elastic arteries in invertebrates: mechanics of the octopus aorta. Science. 1981 Aug 14; 213(4509):759-61. PMID: 7256277.
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